The Grünbach Formation is an Austrian geological formation that dates to the early Campanian age of the Late Cretaceous. it forms part of the Gosau Group, and represents a marine regression event, representing a coastal/brackish environment, being underlain by the marine carbonate Maiersdorf Formation and overlain by the deep marine siliciclastic Piesting Formation. The main lithology is clay, marl, siltstone and sandstone, with a minor conglomerate component. Coal seams have also been noted.[1][2] It is notable for its fossils including those of dinosaurs and plants.
Vertebrate paleofauna
All remains have been found at a single locality, which is a thin marl seam in the Konstantin mining tunnel, within the Felbering Mine in the Neue Welt area north west of Muthmannsdorf in Lower Austria.[3] The initial remains were discovered in 1859 after an ornithopod tooth was found in a piece of coal in a dump outside the mine by Professor Ferdinand Stoliczka, and the productive seam discovered thereafter. The first material was described by Emanuel Bunzel in 1871[4] and then additional material was described by Harry Seeley in 1881.[5] Due to mining activity in the area ceasing at the end of the 19th century, no additional remains have been recovered since.
Squamates
| Squamates of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images | 
| Araeosaurus | A. gracilis | Muthmannsdorf |  | Vertebra[5] |  |  | 
| Crocodyliformes of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images | 
| Doratodon | D. carcharidens | Muthmannsdorf |  | "incomplete mandible, a fragmentary right maxilla, a parietal fragment, and isolated teeth"[6] |  |  | 
| Eusuchia | Indeterminate | Muthmannsdorf |  | "mandible fragment and some postcranial material" |  |  | 
Choristoderes
| Choristoderes of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images | 
| Choristodera | Indeterminate | Muthmannsdorf |  | "Two platycoelous vertebral centra"[7] |  |  | 
Turtles
| Turtles of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images | 
| Dortokidae[8] | Indeterminate | Muthmannsdorf |  | Shell fragments |  |  | 
| cf. Kallokibotioninae | Indeterminate | Muthmannsdorf |  | Shell fragments |  |  | 
Pterosaurs
| Pterosaurs of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images | 
| "Ornithocheirus" | "O". buenzeli | Muthmannsdorf |  |  | Remains found not diagnostic, remains referred to Azdarchidae and Pterodactyloidea indet. |  | 
| Azdarchidae | Indeterminate | Muthmannsdorf |  | "Proximal portion of a humerus"[9] | Previously referred to "O". buenzeli |  | 
| Pterodactyloidea | Indeterminate | Muthmannsdorf |  | "Articular region of a lower jaw, crushed phalangeal fragments" | Previously referred to "O". buenzeli |  | 
Non-avian dinosaurs
| Dinosaurs of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images | 
| Mochlodon | M. suessi | Muthmannsdorf |  | Cranial and postcranial elements |  |  | 
| Struthiosaurus | S. austriacus | Muthmannsdorf |  | Cranial and postcranial elements |  |  | 
| Tetanurae | Indeterminate | Muthmannsdorf |  | "Two fragmentary teeth" | Referred to by the dubious name ‘Megalosaurus pannoniensis’[5] Nearly identical to teeth known from the Csehbánya Formation[10] |  | 
Flora
Most of these specimens were recovered from mining dumps near Grünbach am Schneeberg in lower Austria. The flora of the formation is considered to represent that of a high humidity subtropical climate, typical of the Euro-Sinian phytogeographical region.[11][12][13]
|  |  |  | 
| Color key 
 |  | Notes Uncertain or tentative taxa are in small text;
 crossed outtaxa are discredited. | 
Bryophytes
| Bryophytes of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images | 
| Stereisporites | Indeterminate |  |  |  | Spore, representative of Sphagnaceae |  | 
Lycopodiophyta
| Lycopodiophytes of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images | 
| Camarozonosporites | Indeterminate |  |  |  | Spore, representative of Lycopodiaceae |  | 
| Retitriletes | Indeterminate |  |  |  | Spore, representative of Lycopodium |  | 
Pteridophytes
| Pteridophytes of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images | 
| Appendicisporites | A. tricuspidatus |  |  |  | Spore, probable affinities with Schizaeaceae |  | 
| Cicatricosisporites | Indeterminate |  |  |  | Spore, probable affinities with Schizaeaceae |  | 
| Cingulatisporites | Indeterminate |  |  |  | Spore, affinities with Anemia |  | 
| Cladophlebis | C. gosauensis |  |  |  |  |  | 
| Coniopteris | ? |  |  |  |  |  | 
| Cyathidites | Indeterminate |  |  |  | Spore |  | 
| Echinatisporites | Indeterminate |  |  |  | Spore |  | 
| Equisetites | Indeterminate |  |  |  | A horsetail |  | 
| Gosauopteris | G. danaeoides |  |  |  | Probable affinities with Marattiaceae |  | 
| Gleicheniidites | G. senonicus |  |  |  | Spore |  | 
| Leiotriletes | Indeterminate |  |  |  | Spore, representative of Lygodium |  | 
| Marsileaceaephyllum | M. campanica |  |  |  | Marsileaceae, very similar to extant Marsilea |  | 
| Matonisporites | Indeterminate |  |  |  | Spore,  Found in situ with Monheimia |  | 
| Microphyllopteris | M. austriaca |  |  |  |  |  | 
| Microtaenia | M. austriaca |  |  |  |  |  | 
| Monheimia | M.  ungerii |  |  |  | affinities with Matoniaceae, particularly Phanerosorus |  | 
| Polypodiaceoisporites | Indeterminate |  |  |  | Spore |  | 
| Raphaelia | R. lobifolia |  |  |  |  |  | 
| Sphenopteris | S. gruenbachiana, S. ungeri |  |  |  | Probable affinities with Schizaeaceae |  | 
Gymnosperms
| Gymnosperms of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images | 
| Geinitzia | G. formosa, G. reichenbachii |  |  |  | Pinopsida, Taxodioid affinity |  | 
| cf. Lindleycladus | L. lanceolatus |  |  |  |  |  | 
| Nilsonia | N. cf. holyi |  |  |  | Bennettitales |  | 
| Pagiophyllum | Indeterminate |  |  |  |  |  | 
| Pityosporites | Indeterminate |  |  |  | Pollen |  | 
| Podozamites | Indeterminate |  |  |  |  |  | 
| Taxiodiaceaepollenites | Indeterminate |  |  |  | Pollen, at least 2 taxa |  | 
Angiosperms
Monocots
| Monocots of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images | 
| Arecipites | Indeterminate |  |  |  | Pollen, representative of Palmae |  | 
| Lysichiton | L. austriacus |  |  |  | Araceae, genus extant |  | 
| Gruenbachia | G. pandanoides |  |  |  | Pandanaceae, reproductive organs of Pandanites |  | 
| Pandanites | P. trinervis |  |  |  | Pandanaceae |  | 
| Sabalites | S. longirhachis |  |  |  | Palm |  | 
| Spinizonocolpites | Indeterminate |  |  |  | Pollen, representative of extant Nypa palm |  | 
| Theiaiphyllum | T. kollmannii |  |  |  |  |  | 
Dicots
| Dicots of the Grünbach Formation | 
| Genus | Species | Location | Stratigraphic Position | Material | Notes | Images | 
| Brasenites | B. krasserii |  |  |  | affinity with Brasenia |  | 
| Celastrophyllum | Indeterminate, C. johannae |  |  |  | Hamamelidae |  | 
| Ceratoxylon | Indeterminate |  |  |  | Reproductive structure |  | 
| Clavatipollenites | Indeterminate |  |  |  | Pollen, representative of Chloranthaceae |  | 
| Complexiopollis | Indeterminate |  |  |  | Pollen, affinities with Myricaceae |  | 
| Compositiphyllum | C. serratum |  |  |  |  |  | 
| Debeya | D. insignis |  |  |  |  |  | 
| Dicotylophyllum | Indeterminate |  |  |  |  |  | 
| Ettingshausenia | E. gruenbachiana,  E. cf. laevis, Indeterminate |  |  |  | Platanaceae |  | 
| Extremipollis | Indeterminate |  |  |  | Pollen |  | 
| Grebenkia | G. europeica |  |  |  |  |  | 
| cf. Grevilleophyllum | G. constans |  |  |  |  |  | 
| Hungaropollis | Indeterminate |  |  |  | Pollen |  | 
| Juglandiphyllites | J. pelagicus |  |  |  | possible affinity with Juglandaceae |  | 
| Krutzschipollites | Indeterminate |  |  |  | Pollen |  | 
| Laudaypollis | Indeterminate |  |  |  | Pollen |  | 
| Leguminosites | L. mucronata |  |  |  |  |  | 
| Longanulipollis | Indeterminate |  |  |  | Pollen |  | 
| Menispermites | M. ettingshausenii, M. summesbergerii |  |  |  |  |  | 
| Momipetes | Indeterminate |  |  |  | Pollen, similar to extant Engelhardtia |  | 
| Myricophyllum | M. cf. zenkeri, M. serratum |  |  |  | Affinities with Myrica |  | 
| Myrtacaedites | Indeterminate |  |  |  | Pollen, Representative of Myrtaceae |  | 
| Normapolles | Indeterminate |  |  |  | Pollen, very abundant. possibly associated with Juglandaceae |  | 
| Oculipollis | O. sp, O. parvoculus O.zaklinskaiaiae |  |  |  | Pollen |  | 
| cf. Pandemophyllum | Indeterminate, P. cf. proteoides |  |  |  |  |  | 
| Pseudoplicapollis | Indeterminate |  |  |  | Pollen |  | 
| Quereuxia | Q. angulata |  |  |  | Possible affinity with Trapa |  | 
| Rodgersia | Indeterminate |  |  |  |  |  | 
| Semioculipollis | Indeterminate |  |  |  | Pollen |  | 
| Suemegipollis | Indeterminate |  |  |  | Pollen |  | 
| Ternstroemites | T. (?) neueweltensis |  |  |  |  |  | 
| Tricolpopollenites | Indeterminate |  |  |  | Pollen, At least 8 distinct taxa |  | 
| Viburniphyllum | V. ermannii, V. austriacum |  |  |  |  |  | 
References
- ^ Draganits, Erich; Wagreich, Michael; Hofer, Gerald; Hofmann, Christa-Charlotte; Reischenbacher, Doris; Grundtner, Marie-Louise; Neuhuber, Stephanie; Bottig, Magda (2011). "Stratigraphy and geochemical characterisation of Upper Cretaceous non-marine - marine cycles (Grünbach Formation, Gosau Group, Austria". Austrian Journal of Earth Sciences. 104 (2): 90–107.
- ^ Grundtner, Marie-Louise (2011). Sedimentologie und Sedimentpetrographie der Gosau-Gruppe bei Dreistetten-Muthmannsdorf (Niederösterreichische Kalkalpen) (Master's thesis). University of Vienna. pp. 21–35, 51–57, 71. doi:10.25365/thesis.17461.
- ^ Sachs, Sven; Hornung, Jahn J. (May 2006). "Juvenile ornithopod (Dinosauria: Rhabdodontidae) remains from the Upper Cretaceous (Lower Campanian, Gosau Group) of Muthmannsdorf (Lower Austria)". Geobios. 39 (3): 415–425. doi:10.1016/j.geobios.2005.01.003. ISSN 0016-6995.
- ^ Bunzel, Emanuel (1871). "Die Reptilfauna der Gosau-Formation in der Neuen Welt bei Wiener-Neustadt". Abhandlungen der Kaiserlich-königlichen Geologischen Reichsanstalt. 5: 1–18.
- ^ a b c Seeley, H. G. (1881-02-01). "The Reptile Fauna of the Gosau Formation preserved in the Geological Museum of the University of Vienna: With a Note on the Geological Horizon of the Fossils at Neue Welt, west of Wiener Neustadt, by Edw. Suess, Ph.D., F.M.G.S., &c., Professor of Geology in the University of Vienna, &c". Quarterly Journal of the Geological Society. 37 (1–4): 620–706. doi:10.1144/GSL.JGS.1881.037.01-04.49. ISSN 0370-291X. S2CID 219235284.
- ^ Buffetaut, Eric (1979). "Revision der Crocodylia (Reptilia) aus den Gosau-Schichten (Ober-Kreide) von Österreich". Beiträge zur Paläontologie von Österreich. 6: 89–105.
- ^ Buffetaut, Eric (1989). "Erster nachweis von Choristodera (Reptilia, Diapsida) in der Oberkreide Europas: Champsosaurierwirbel aus den Gosau-Schichten (Campan) Niederösterreichs". Sitzungsberichten der Österreichs Akademis der Wissenschaften Mathematisch-Naturwissenschaftlichen Klasse, Abteilung. 197: 389–394.
- ^ Rabi, Márton; Vremir, Mátyás; Tong, Haiyan (2012-09-01), "Preliminary Overview of Late Cretaceous Turtle Diversity in Eastern Central Europe (Austria, Hungary, and Romania)", Vertebrate Paleobiology and Paleoanthropology, Springer Netherlands, pp. 307–336, doi:10.1007/978-94-007-4309-0_19, ISBN 9789400743083
- ^ Buffetaut, Eric; Ősi, Attila; Prondvai, Edina (March 2011). "The pterosaurian remains from the Grünbach Formation (Campanian, Gosau Group) of Austria: a reappraisal of 'Ornithocheirus buenzeli'". Geological Magazine. 148 (2): 334–339. Bibcode:2011GeoM..148..334B. doi:10.1017/S0016756810000981. ISSN 1469-5081. S2CID 131376979.
- ^ Csiki-Sava, Zoltan; Buffetaut, Eric; Ősi, Attila; Pereda-Suberbiola, Xabier; Brusatte, Stephen L. (2015-01-08). "Island life in the Cretaceous - faunal composition, biogeography, evolution, and extinction of land-living vertebrates on the Late Cretaceous European archipelago". ZooKeys (469): 1–161. doi:10.3897/zookeys.469.8439. ISSN 1313-2970. PMC 4296572. PMID 25610343.
- ^ Herman, Alexei B.; KVAČEK, JIŘÍ (2007). "Early Campanian Grünbach flora of Austria: systematic composition and palaeoclimatic interpretations". Acta Palaeobotanica. 47 (1): 37–55.
- ^ KVACEK, JIRÍ; HERMAN, ALEXEI (2004). "The Campanian Grünbach Flora of Lower Austria: palaeoecological interpretations". Annalen des Naturhistorischen Museums in Wien. 106 A.
- ^ B., Herman, Alexei (2010). Late Cretaceous Grünbach Flora of Austria. Kvaček, Jiří. Wien: Naturhistorisches Museum. ISBN 9783902421432. OCLC 517260675.{{cite book}}:  CS1 maint: multiple names: authors list (link)